*, *::before, *::after { box-sizing: border-box; margin: 0; padding: 0; }

:root {
  --canvas: #f7f7f4;
  --canvas-soft: #fafaf7;
  --surface-card: #ffffff;
  --surface-strong: #e6e5e0;
  --hairline: #e6e5e0;
  --hairline-soft: #efeee8;
  --hairline-strong: #cfcdc4;
  --ink: #26251e;
  --body: #5a5852;
  --body-strong: #26251e;
  --muted: #807d72;
  --muted-soft: #a09c92;
  --primary: #f54e00;
  --primary-active: #d04200;
  --on-primary: #ffffff;
  --success: #1f8a65;
  --error: #cf2d56;
  --font-sans: 'Inter', system-ui, 'Helvetica Neue', Helvetica, Arial, sans-serif;
  --font-mono: 'JetBrains Mono', monospace;
  --radius-xs: 4px;
  --radius-sm: 6px;
  --radius-md: 8px;
  --radius-lg: 12px;
  --radius-xl: 16px;
  --radius-pill: 9999px;
}

html { scroll-behavior: smooth; }

body {
  font-family: var(--font-sans);
  font-size: 16px;
  font-weight: 400;
  line-height: 1.5;
  color: var(--body);
  background-color: var(--canvas);
  -webkit-font-smoothing: antialiased;
}

a { color: var(--ink); text-decoration: none; }
a:hover { color: var(--primary); }
img { max-width: 100%; height: auto; display: block; }
ul { list-style: none; }

.container {
  max-width: 1200px;
  margin: 0 auto;
  padding: 0 24px;
}

.site-nav {
  background: var(--canvas);
  border-bottom: 1px solid var(--hairline);
  position: sticky;
  top: 0;
  z-index: 100;
  height: 64px;
}

.nav-inner {
  max-width: 1200px;
  margin: 0 auto;
  padding: 0 24px;
  display: flex;
  align-items: center;
  justify-content: space-between;
  height: 64px;
}

.nav-logo {
  font-size: 18px;
  font-weight: 600;
  color: var(--ink);
  letter-spacing: -0.3px;
}

.nav-menu {
  display: flex;
  align-items: center;
  gap: 32px;
}

.nav-menu li a {
  font-size: 14px;
  font-weight: 500;
  color: var(--body);
  line-height: 1.4;
  transition: color 0.15s;
}

.nav-menu li a:hover { color: var(--ink); }

.nav-toggle {
  display: none;
  flex-direction: column;
  gap: 5px;
  background: none;
  border: none;
  cursor: pointer;
  padding: 4px;
}

.nav-toggle span {
  display: block;
  width: 22px;
  height: 2px;
  background: var(--ink);
  border-radius: 2px;
  transition: all 0.2s;
}

.hero {
  padding: 80px 0;
  border-bottom: 1px solid var(--hairline);
}

.hero-inner {
  max-width: 1200px;
  margin: 0 auto;
  padding: 0 24px;
}

.hero-label {
  display: inline-block;
  font-size: 11px;
  font-weight: 600;
  letter-spacing: 0.88px;
  text-transform: uppercase;
  background: var(--surface-strong);
  color: var(--ink);
  border-radius: var(--radius-pill);
  padding: 4px 10px;
  margin-bottom: 24px;
}

.hero h1 {
  font-size: 56px;
  font-weight: 400;
  line-height: 1.1;
  letter-spacing: -1.5px;
  color: var(--ink);
  max-width: 780px;
  margin-bottom: 20px;
}

.hero-sub {
  font-size: 18px;
  line-height: 1.55;
  color: var(--body);
  max-width: 600px;
  margin-bottom: 40px;
}

.hero-img {
  margin-top: 48px;
  border-radius: var(--radius-lg);
  border: 1px solid var(--hairline);
  overflow: hidden;
  max-height: 480px;
}

.hero-img img {
  width: 100%;
  height: 480px;
  object-fit: cover;
}

.section {
  padding: 80px 0;
  border-bottom: 1px solid var(--hairline-soft);
}

.section-label {
  font-size: 11px;
  font-weight: 600;
  letter-spacing: 0.88px;
  text-transform: uppercase;
  color: var(--muted);
  margin-bottom: 12px;
}

.section-title {
  font-size: 36px;
  font-weight: 400;
  line-height: 1.2;
  letter-spacing: -0.72px;
  color: var(--ink);
  margin-bottom: 16px;
}

.section-sub {
  font-size: 16px;
  line-height: 1.5;
  color: var(--body);
  max-width: 640px;
  margin-bottom: 40px;
}

.card-grid {
  display: grid;
  grid-template-columns: repeat(3, 1fr);
  gap: 24px;
}

.card {
  background: var(--surface-card);
  border: 1px solid var(--hairline);
  border-radius: var(--radius-lg);
  padding: 24px;
}

.card-img {
  border-radius: var(--radius-md);
  overflow: hidden;
  margin-bottom: 20px;
}

.card-img img {
  width: 100%;
  height: 200px;
  object-fit: cover;
}

.card-meta {
  font-size: 11px;
  font-weight: 600;
  letter-spacing: 0.88px;
  text-transform: uppercase;
  color: var(--muted);
  margin-bottom: 8px;
}

.card h3 {
  font-size: 18px;
  font-weight: 600;
  line-height: 1.4;
  color: var(--ink);
  margin-bottom: 10px;
}

.card p {
  font-size: 14px;
  line-height: 1.5;
  color: var(--body);
}

.card-link {
  display: inline-flex;
  align-items: center;
  gap: 6px;
  font-size: 14px;
  font-weight: 500;
  color: var(--ink);
  margin-top: 16px;
  border-bottom: 1px solid var(--hairline-strong);
  padding-bottom: 2px;
}

.card-link:hover { color: var(--primary); border-color: var(--primary); }

.two-col {
  display: grid;
  grid-template-columns: 1fr 1fr;
  gap: 48px;
  align-items: start;
}

.two-col-img {
  border-radius: var(--radius-lg);
  overflow: hidden;
  border: 1px solid var(--hairline);
}

.two-col-img img {
  width: 100%;
  height: 360px;
  object-fit: cover;
}

.two-col-content h2 {
  font-size: 26px;
  font-weight: 400;
  line-height: 1.25;
  letter-spacing: -0.325px;
  color: var(--ink);
  margin-bottom: 16px;
}

.two-col-content p {
  font-size: 16px;
  line-height: 1.6;
  color: var(--body);
  margin-bottom: 16px;
}

.info-list {
  margin-top: 24px;
  border-top: 1px solid var(--hairline);
}

.info-list li {
  display: flex;
  gap: 12px;
  padding: 12px 0;
  border-bottom: 1px solid var(--hairline-soft);
  font-size: 14px;
  color: var(--body);
}

.info-list li strong {
  color: var(--ink);
  min-width: 140px;
  font-weight: 600;
}

.btn-primary {
  display: inline-block;
  background: var(--primary);
  color: var(--on-primary);
  font-size: 14px;
  font-weight: 500;
  line-height: 1;
  padding: 10px 18px;
  height: 40px;
  border-radius: var(--radius-md);
  border: none;
  cursor: pointer;
  transition: background 0.15s;
  text-decoration: none;
}

.btn-primary:hover { background: var(--primary-active); color: var(--on-primary); }

.btn-secondary {
  display: inline-block;
  background: var(--surface-card);
  color: var(--ink);
  font-size: 14px;
  font-weight: 500;
  line-height: 1;
  padding: 10px 18px;
  height: 40px;
  border-radius: var(--radius-md);
  border: 1px solid var(--hairline-strong);
  cursor: pointer;
  transition: border-color 0.15s;
  text-decoration: none;
}

.btn-secondary:hover { border-color: var(--ink); color: var(--ink); }

.article-header {
  padding: 64px 0 48px;
  border-bottom: 1px solid var(--hairline);
}

.article-header .breadcrumb {
  font-size: 13px;
  color: var(--muted);
  margin-bottom: 20px;
}

.article-header .breadcrumb a { color: var(--muted); }
.article-header .breadcrumb a:hover { color: var(--ink); }

.article-header h1 {
  font-size: 40px;
  font-weight: 400;
  line-height: 1.15;
  letter-spacing: -0.8px;
  color: var(--ink);
  max-width: 800px;
  margin-bottom: 16px;
}

.article-header .article-desc {
  font-size: 18px;
  line-height: 1.55;
  color: var(--body);
  max-width: 680px;
  margin-bottom: 20px;
}

.article-meta {
  font-size: 13px;
  color: var(--muted);
}

.article-body {
  padding: 48px 0 80px;
}

.article-layout {
  display: grid;
  grid-template-columns: 1fr 300px;
  gap: 64px;
  max-width: 1200px;
  margin: 0 auto;
  padding: 0 24px;
}

.article-content h2 {
  font-size: 26px;
  font-weight: 400;
  line-height: 1.25;
  letter-spacing: -0.325px;
  color: var(--ink);
  margin: 40px 0 16px;
}

.article-content h2:first-child { margin-top: 0; }

.article-content h3 {
  font-size: 18px;
  font-weight: 600;
  line-height: 1.4;
  color: var(--ink);
  margin: 28px 0 12px;
}

.article-content p {
  font-size: 16px;
  line-height: 1.65;
  color: var(--body);
  margin-bottom: 20px;
}

.article-content ul, .article-content ol {
  margin: 0 0 20px 20px;
  list-style: disc;
}

.article-content ol { list-style: decimal; }

.article-content li {
  font-size: 16px;
  line-height: 1.65;
  color: var(--body);
  margin-bottom: 6px;
}

.article-content .article-img {
  margin: 32px 0;
  border-radius: var(--radius-lg);
  overflow: hidden;
  border: 1px solid var(--hairline);
}

.article-content .article-img img {
  width: 100%;
  height: 360px;
  object-fit: cover;
}

.article-content .img-caption {
  font-size: 13px;
  color: var(--muted);
  padding: 8px 12px;
  border-top: 1px solid var(--hairline-soft);
  background: var(--canvas-soft);
}

.article-content .data-table {
  width: 100%;
  border-collapse: collapse;
  margin: 24px 0;
  font-size: 14px;
}

.article-content .data-table th {
  background: var(--surface-strong);
  color: var(--ink);
  font-weight: 600;
  padding: 10px 14px;
  text-align: left;
  border: 1px solid var(--hairline);
}

.article-content .data-table td {
  padding: 10px 14px;
  border: 1px solid var(--hairline-soft);
  color: var(--body);
  background: var(--surface-card);
}

.article-content .data-table tr:nth-child(even) td {
  background: var(--canvas-soft);
}

.article-content .highlight-box {
  background: var(--surface-card);
  border: 1px solid var(--hairline);
  border-left: 3px solid var(--ink);
  border-radius: var(--radius-md);
  padding: 20px 24px;
  margin: 28px 0;
}

.article-content .highlight-box p {
  margin: 0;
  font-size: 15px;
  color: var(--body);
}

.article-sidebar .sidebar-card {
  background: var(--surface-card);
  border: 1px solid var(--hairline);
  border-radius: var(--radius-lg);
  padding: 24px;
  margin-bottom: 24px;
}

.article-sidebar .sidebar-card h4 {
  font-size: 13px;
  font-weight: 600;
  letter-spacing: 0.88px;
  text-transform: uppercase;
  color: var(--muted);
  margin-bottom: 14px;
}

.article-sidebar .sidebar-card ul {
  margin: 0;
}

.article-sidebar .sidebar-card ul li {
  padding: 6px 0;
  border-bottom: 1px solid var(--hairline-soft);
  font-size: 14px;
}

.article-sidebar .sidebar-card ul li:last-child { border-bottom: none; }

.article-sidebar .sidebar-card ul li a {
  color: var(--body);
  line-height: 1.4;
}

.article-sidebar .sidebar-card ul li a:hover { color: var(--ink); }

.contact-form-section {
  background: var(--surface-card);
  border: 1px solid var(--hairline);
  border-radius: var(--radius-lg);
  padding: 32px;
  margin-top: 48px;
}

.contact-form-section h3 {
  font-size: 22px;
  font-weight: 400;
  letter-spacing: -0.11px;
  color: var(--ink);
  margin-bottom: 8px;
}

.contact-form-section .form-desc {
  font-size: 15px;
  color: var(--body);
  margin-bottom: 24px;
}

.form-row {
  display: grid;
  grid-template-columns: 1fr 1fr;
  gap: 16px;
  margin-bottom: 16px;
}

.form-group {
  margin-bottom: 16px;
}

.form-group label {
  display: block;
  font-size: 13px;
  font-weight: 600;
  color: var(--ink);
  margin-bottom: 6px;
}

.form-group input,
.form-group textarea,
.form-group select {
  width: 100%;
  background: var(--surface-card);
  color: var(--ink);
  border: 1px solid var(--hairline-strong);
  border-radius: var(--radius-md);
  padding: 12px 16px;
  height: 44px;
  font-size: 16px;
  font-family: var(--font-sans);
  transition: border-color 0.15s;
  outline: none;
}

.form-group textarea {
  height: auto;
  min-height: 120px;
  resize: vertical;
}

.form-group input:focus,
.form-group textarea:focus {
  border-color: var(--ink);
}

.form-group input.error,
.form-group textarea.error {
  border-color: var(--error);
}

.form-error {
  font-size: 13px;
  color: var(--error);
  margin-top: 4px;
  display: none;
}

.form-error.visible { display: block; }

.form-message {
  padding: 14px 18px;
  border-radius: var(--radius-md);
  font-size: 15px;
  display: none;
  margin-top: 16px;
}

.form-message.success {
  background: #e8f5f0;
  color: var(--success);
  border: 1px solid #b8ddd2;
  display: block;
}

.page-header {
  padding: 64px 0 48px;
  border-bottom: 1px solid var(--hairline);
}

.page-header h1 {
  font-size: 40px;
  font-weight: 400;
  line-height: 1.15;
  letter-spacing: -0.8px;
  color: var(--ink);
  margin-bottom: 16px;
}

.page-header p {
  font-size: 18px;
  line-height: 1.55;
  color: var(--body);
  max-width: 640px;
}

.page-content {
  padding: 48px 0 80px;
  max-width: 800px;
}

.page-content h2 {
  font-size: 22px;
  font-weight: 400;
  letter-spacing: -0.11px;
  color: var(--ink);
  margin: 36px 0 12px;
}

.page-content h2:first-child { margin-top: 0; }

.page-content p {
  font-size: 16px;
  line-height: 1.65;
  color: var(--body);
  margin-bottom: 16px;
}

.page-content ul {
  margin: 0 0 16px 20px;
  list-style: disc;
}

.page-content li {
  font-size: 16px;
  line-height: 1.65;
  color: var(--body);
  margin-bottom: 4px;
}

.page-updated {
  font-size: 13px;
  color: var(--muted);
  margin-top: 40px;
  padding-top: 20px;
  border-top: 1px solid var(--hairline-soft);
}

.species-grid {
  display: grid;
  grid-template-columns: repeat(2, 1fr);
  gap: 20px;
  margin: 32px 0;
}

.species-card {
  background: var(--surface-card);
  border: 1px solid var(--hairline);
  border-radius: var(--radius-lg);
  padding: 20px;
}

.species-card .sci-name {
  font-size: 13px;
  font-style: italic;
  color: var(--muted);
  margin-bottom: 4px;
}

.species-card h4 {
  font-size: 16px;
  font-weight: 600;
  color: var(--ink);
  margin-bottom: 8px;
}

.species-card p {
  font-size: 14px;
  line-height: 1.5;
  color: var(--body);
  margin: 0;
}

.site-footer {
  background: var(--canvas);
  border-top: 1px solid var(--hairline);
  padding: 64px 0 0;
}

.footer-inner {
  max-width: 1200px;
  margin: 0 auto;
  padding: 0 24px;
  display: grid;
  grid-template-columns: 2fr 1fr 1fr 1fr;
  gap: 48px;
  padding-bottom: 48px;
  border-bottom: 1px solid var(--hairline-soft);
}

.footer-brand {
  font-size: 16px;
  font-weight: 600;
  color: var(--ink);
  display: block;
  margin-bottom: 12px;
}

.footer-col p {
  font-size: 14px;
  line-height: 1.6;
  color: var(--body);
  margin-top: 8px;
}

.footer-col h3 {
  font-size: 13px;
  font-weight: 600;
  letter-spacing: 0.88px;
  text-transform: uppercase;
  color: var(--muted);
  margin-bottom: 14px;
}

.footer-col ul li {
  margin-bottom: 8px;
}

.footer-col ul li a {
  font-size: 14px;
  color: var(--body);
}

.footer-col ul li a:hover { color: var(--ink); }

.footer-bottom {
  max-width: 1200px;
  margin: 0 auto;
  padding: 20px 24px;
  font-size: 13px;
  color: var(--muted);
}

.footer-bottom a { color: var(--muted); }
.footer-bottom a:hover { color: var(--ink); }
.footer-bottom p { margin-bottom: 4px; }
.footer-disclaimer { font-size: 12px; color: var(--muted-soft); }

.cookie-banner {
  position: fixed;
  bottom: 0;
  left: 0;
  right: 0;
  background: var(--ink);
  color: var(--canvas);
  padding: 16px 24px;
  z-index: 200;
  display: none;
}

.cookie-banner.visible { display: block; }

.cookie-inner {
  max-width: 1200px;
  margin: 0 auto;
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: 24px;
  flex-wrap: wrap;
}

.cookie-inner p {
  font-size: 14px;
  color: #d4d3cc;
  flex: 1;
}

.cookie-actions {
  display: flex;
  gap: 10px;
  flex-shrink: 0;
}

@media (max-width: 1024px) {
  .card-grid { grid-template-columns: repeat(2, 1fr); }
  .footer-inner { grid-template-columns: 1fr 1fr; }
  .article-layout { grid-template-columns: 1fr; }
  .article-sidebar { display: grid; grid-template-columns: 1fr 1fr; gap: 20px; }
}

@media (max-width: 768px) {
  .hero h1 { font-size: 36px; letter-spacing: -1px; }
  .section-title { font-size: 26px; }
  .two-col { grid-template-columns: 1fr; }
  .nav-menu {
    display: none;
    position: absolute;
    top: 64px;
    left: 0;
    right: 0;
    background: var(--canvas);
    border-bottom: 1px solid var(--hairline);
    flex-direction: column;
    align-items: flex-start;
    padding: 16px 24px;
    gap: 0;
  }
  .nav-menu.open { display: flex; }
  .nav-menu li { width: 100%; }
  .nav-menu li a { display: block; padding: 10px 0; border-bottom: 1px solid var(--hairline-soft); }
  .nav-toggle { display: flex; }
  .form-row { grid-template-columns: 1fr; }
  .species-grid { grid-template-columns: 1fr; }
  .article-sidebar { grid-template-columns: 1fr; }
}

@media (max-width: 640px) {
  .card-grid { grid-template-columns: 1fr; }
  .footer-inner { grid-template-columns: 1fr; gap: 32px; }
  .hero { padding: 48px 0; }
  .section { padding: 48px 0; }
  .article-header h1 { font-size: 28px; }
}